Biggest Loser from last crash

Now that we seem to be on a run up to a bull run, let's remember to be wary and careful of scams. And to drive the point home, in your opinion of all the losses from the last crash, who do you think on average had the biggest losses, are they LUNA/UST investors, Celsius, FTX, among others? Can you please elaborate why you think so, this will help newbies and even veteran crypto investors alike.

If I understand it, the catalyst really was the crash of LUNA depegging UST, which was itself brought on by an investor shorting LUNA because they saw the flaw in its innards

(Edit: please exclude the perpetrators themselves)